Paarthurnax frozen in mid-air

[Yatzstar]

Basically, Paarthurnax is frozen in what I would assume is the dragon equivalent of a t-pose. Level flight, wing spread, but frozen in place. He still moves his neck and head to roar, and I can still talk to him, but he'll only move upon entering combat, after which he flies off into the distance, does not return to sit on his wall, and goes back to the t-pose after quitting the game. He also doesn't respond to Dragonrend (he just displays the in-flight 'stagger' animation and goes back to t-pose) or any other type of attack. I've tried disabling and re-enabling him, resetai, and even spawning a new copy of him (which stands on the ground in the default fashion, but upon entering and exiting combat flies off like the original) but none of them worked.

 

For reference, the mods I use that affect dragons in any way are these:

KS Dragon Overhaul

Dragon Animation Replacer (SE)

Diverse Dragons Collection SE

Bellyache's HD Dragon Replacer Pack (SE)

Splendor - Dragon Variants SE

The Paarthurnax Dilemma

 

I've uninstalled Dragon Animation Replacer, KS Dragon Overhaul and Bellyache's HD Dragon Replacer, but nothing changes. I'm still working on uninstalling DDC (my SKSE64 is iffy at best when it comes to launching the game with it) but I'm hoping to get that uninstalled at some point. It's hard to pinpoint when exactly the t-pose started, but I'm fairly certain it was after the Throat of the World fight with Alduin, given how combat seems to affect things. (If it affects anything, I've proceeded in the main quest up until the 'Talk to Odahviing' stage of The World-Eater's Eyrie)

 

I can post my entire load order if need be. Any help would be greatly appreciated--I just want my bro to sit on his wall again like normal.